Indices |
Formulas |
References |
meaning |
species richness |
SR = (S – 1) / ln(N)
Where,
SR = Species richness index
S = Total number of species
N = Total number of individuals in the sample |
Margalef [50] |
A measure for species diversity, the higher the index the greater the diversity |
diversity index |
where, Pi = ni /N
Pi = the proportion of individuals found for the ith species
ni = the number of individuals of the ith species
N = the total number of individuals of all species |
Shannon and Wiener [51] |
A measure for species diversity taking into account how evenly the individuals are distributed among the species, for a given number of species, the index is maximized when all species are equally abundant |
dominance |

Where,
ID = Index of dominance
S = Number of species
ni = Number of individuals of the ith species
N = Total number of individuals of all species |
Simpson [52] |
A measure for the degree of concentration of species, the index obtains small values in datasets of high diversity and large values in datasets of low diversity |
evenness |

Where,
E = Species evenness index
H = Shannon-Wiener Index of diversity
S = Total no. of species |
Pileou [53] |
A measure to quantify how equal a community is in numbers of each species, a high value indicates a high evenness |
similarity |
IS = (3×N) / (C+D+S)
Where,
IS = Index of Similarity
N = number of common species in the sites
C = Species present in site C
D = Species present in site D
S = Species present in site S |
Sørensen [54] |
A measure for comparing the similarity of different sites with regard to common species, a high value indicates large similarity between the sites |